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The main proteins responsible are sulfhydryl oxidases, which often incorporate iron, copper, or flavin binding domains to participate in oxidation-reduction reactions (Kodali and Thorpe, 2010)

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2020) Car-OO · may behave as a prooxidant by further reaction with unsaturated lipids Thus carotenoids behave under normal conditions as antioxidants, however, under a high partial pressure of oxygen (and high concentration) they may lose their antioxidant properties and behave as prooxidants